1. Department of Animal and Plant Sciences; University of Sheffield; Sheffield S10 2TN UK
2. Evolutionary Ecology; Max Planck Institute for Evolutionary Biology; 24306 Ploen Germany
3. The University of Queensland; Queensland Brain Institute; Brisbane QLD 4072 Australia
4. Livestock Industries; CSIRO 4067; Brisbane QLD Australia
5. The University of Queensland Diamantina Institute; Brisbane QLD 4072 Australia
6. Institute of Evolutionary Biology; School of Biological Sciences; University of Edinburgh; Edinburgh EH9 3JT UK