1. Yunnan Key Laboratory of Plant Reproductive Adaptation and Evolutionary Ecology Yunnan University Kunming 650500 China
2. Laboratory of Ecology and Evolutionary Biology, School of Ecology and Environmental Sciences Yunnan University Kunming 650091 China
3. Department of Ecology and Evolutionary Biology University of Toronto 25 Willcocks Street Toronto Ontario M5S 3B2 Canada
4. Kunming Institute of Botany Chinese Academy of Sciences Kunming 650201 China