1. School of Biology; Newcastle University; Newcastle upon Tyne NE1 7RU UK
2. Biosciences Division; Bioenergy Science Center; Oak Ridge National Laboratory; Oak Ridge TN 37831-6407 USA
3. Climate Change Science Institute; Environmental Sciences Division; Oak Ridge National Laboratory; Oak Ridge TN 37831-6301 USA
4. Department of Plant Sciences; Institute of Integrative Biology; University of Liverpool; Liverpool L69 7ZB UK
5. Department of Biochemistry and Molecular Biology, MS330; University of Nevada; Reno NV 89557-0330 USA