1. Institute of Mountain Hazards and Environment; Chinese Academy of Sciences; Chengdu 610041 China
2. Sichuan Province Key Laboratory of Ecological Security and Protection; Mianyang Normal University; Mianyang 621000 China
3. Chengdu Institute of Biology; Chinese Academy of Sciences; Chengdu 610041 China
4. University of Chinese Academy of Sciences; Beijing 100039 China
5. Department of Agricultural Sciences; University of Helsinki; Helsinki FI-00014 Finland
6. Department of Plant Physiology; Estonian University of Life Sciences; Tartu 51014 Estonia