1. Lancaster Environment Centre; Lancaster University; Bailrigg, Lancaster LA1 4YQ U.K.
2. Tropical Forest Research Centre; CSIRO; PO Box 780, Atherton Queensland 4883 Australia
3. Present address: Karisoke Research Centre, The Dian Fossey Gorilla Fund International; NR 4 RD 58, B.P. 105, Ruhengeri/Musanze Rwanda
4. Centre for Tropical Environmental and Sustainability Science; College of Marine & Environmental Sciences, James Cook University; Townsville Queensland 4811 Australia
5. Biome5 Pty Ltd.; PO Box 1200, Atherton Queensland 4883 Australia
6. Research Institute for the Environment and Livelihoods; Charles Darwin University; Darwin Northern Territory 0909 Australia