1. Centre d'Ecologie et des Sciences de la Conservation (CESCO UMR7204), Sorbonne Universités, MNHN, CNRS, UPMC, CP51; 55-61 rue Buffon 75005 Paris France
2. Department of Biology; McGill University; 1205 ave Docteur Penfield Montreal Quebec H3A 1B1 Canada
3. African Centre for DNA Barcoding; University of Johannesburg; APK Campus, PO Box 524, Auckland Park 2006 Johannesburg South Africa
4. Department of Biology; University of Toronto Scarborough; 1265 Military Trail Toronto Ontario M1C 1A4 Canada
5. Mathematical Ecology Research Group, Department of Zoology; University of Oxford; South Parks Road Oxford, OX1 3PS U.K.