1. College of Life Sciences; Capital Normal University; 105 Xisanhuanbeilu, Haidian District Beijing 100048 China
2. Department of Entomology; University of Kentucky; S225 Agricultural Science Center North Lexington KY 40546-0091 USA
3. Palaeontological Institute; Russian Academy of Sciences; 123 Profsoyuznayaul. Moscow 117997 Russia
4. Department of Palaeontology; Natural History Museum; Cromwell Road London SW7 5BD UK
5. Department of Entomology; China Agricultural University; No. 2 Yuanmingyuan West Road Beijing 100193 China