1. Department of Plant Protection; College of Plant Science and Technology and the Key Laboratory of Crop Disease Monitoring and Safety Control in Hubei Province, Huazhong Agricultural University; Wuhan 430070 China
2. Department of Agricultural and Environmental Sciences; Clemson University; Clemson SC 29634 USA
3. The Key Laboratory of Horticultural Plant Biology, Ministry of Education, Huazhong Agricultural University; Wuhan 430070 China