1. Department of Biology University of Texas at Arlington Arlington TX USA
2. Amphibian and Reptile Diversity Research Center Department of Biology University of Texas at Arlington Arlington TX USA
3. Department of Biology Indiana University of Pennsylvania Indiana PA USA
4. California Academy of Sciences San Francisco CA USA
5. Centro Zamorano de Biodiversidad Departamento de Ambiente y Desarrollo Escuela Agrícola Panamericana Zamorano Municipalidad de San Antonio de Oriente Francisco Morazán Honduras