1. Romberg Tiburon Center for Environmental Studies; San Francisco State University; 3150 Paradise Drive Tiburon California 94920 USA
2. Department of Biology; Western University; London Ontario Canada N6A 5B7
3. Northwest Fisheries Science Center; National Marine Fisheries Service; National Oceanic and Atmospheric Administration; 2725 Montlake Blvd. E. Seattle Washington 98112 USA