1. Laboratory of Genetics Wageningen University & Research Droevendaalsesteeg 1 Wageningen 6700 AA the Netherlands
2. Department of Developmental Biology, Institut für Pflanzenwissenschaften und Mikrobiologie University of Hamburg Ohnhorststrasse 18 Hamburg 22609 Germany
3. Department of Biological Sciences National University of Singapore 14 Science Drive 4 Singapore 117543 Singapore
4. A*STAR, Institute of Molecular and Cell Biology (IMCB) 61 Biopolis Drive Proteos 138673 Singapore
5. UKE Martinistrasse 52 20251 Hamburg Germany