1. Institute of Arctic and Alpine Research University of Colorado Campus Box 450 Boulder Colorado80309 USA
2. Department of Environmental Sciences Western Washington University Bellingham Washington98225 USA
3. Division of the Biological Sciences The University of Chicago Chicago Illinois60637 USA
4. Scripps Institution of Oceanography University of California La Jolla California USA
5. Department of Microbial and Environmental Genomics J. Craig Venter Institute La Jolla California92037 USA
6. Department of Biology Evolutionary Ecology Laboratories and Monte L. Bean Museum Brigham Young University Provo Utah84602 USA