1. Department of Microbiology University of Tennessee Knoxville Tennessee37996 USA
2. Biology and Paleo Environment Division Lamont‐Doherty Earth Observatory Columbia University Palisades New York10964 USA
3. Department of Earth and Environmental Sciences Columbia University Palisades New York10964 USA
4. School of Marine and Atmospheric Sciences Stony Brook University Stony Brook New York11790 USA