1. Key Laboratory of Plant Development and Environmental Adaption Biology, Ministry of Education, School of Life Science Shandong University Qingdao 266237 P.R. China
2. Key Laboratory of Horticultural Plant Biology, Ministry of Education, College of Horticulture and Forestry Sciences Huazhong Agricultural University Wuhan 430070 P.R. China
3. National Centre for Plant Gene Research (Beijing), Innovation Academy for Seed Design, Institute of Genetics and Developmental Biology Chinese Academy of Sciences Beijing 100101 P.R. China
4. College of Advanced Agricultural Sciences University of Chinese Academy of Sciences Beijing 100039 P.R. China
5. Division of Integrative Biosciences and Biotechnology Pohang University of Science and Technology Pohang 790‐784 South Korea
6. Department of Life Sciences Pohang University of Science and Technology Pohang South Korea