1. Center for Genomics and Biotechnology, Fujian Provincial Key Laboratory of Haixia Applied Plant Systems Biology, Key Laboratory of Genetics, Breeding and Multiple Utilization of Corps Ministry of Education, Fujian Agriculture and Forestry University Fuzhou 350002 Fujian China
2. Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture Wuhan Botanical Garden, Chinese Academy of Sciences Wuhan 430074 China
3. Center of Economic Botany Core Botanical Gardens, Chinese Academy of Sciences Wuhan 430074 China
4. Virginia Cooperative of Extension Virginia Polytechnic Institute and State University Blacksburg VA 24061 USA
5. Department of Plant Biology University of Illinois at Urbana‐Champaign Urbana IL 61801 USA