1. CAS Key Laboratory of Tropical Forest Ecology Xishuangbanna Tropical Botanical Garden Chinese Academy of Sciences Menglun Mengla 666303 China
2. Center of Plant Ecology Core Botanical Gardens Chinese Academy of Sciences Xishuangbanna 666303 China
3. Department of Ecology and Evolutionary Biology University of California Los Angeles Los Angeles CA 90095 USA
4. The James Hutton Institute Aberdeen AB15 8QH UK
5. Institute of Wetland Ecology & Clone Ecology Zhejiang Provincial Key Laboratory of Plant Evolutionary Ecology and Conservation Taizhou University Taizhou 318000 China