1. School of Biomedical Sciences University of Nottingham Medical School Nottingham, NG7 2UH UK
2. School of Physiology and Pharmacology University of Bristol Bristol BS8 1TD UK
3. Neuroscience Division, Medical Education Institute, Ninewells Hospital and Medical School University of Dundee Dundee DD1 9SY UK
4. Centre for Integrative Physiology University of Edinburgh Edinburgh EH8 9XD UK