1. Laboratory of Forest Biology; Division of Forest and Biomaterials Science; Graduate School of Agriculture; Kyoto University; Kyoto Japan
2. Tama Zoo; Hino-shi Tokyo Japan
3. Faculty of Symbiotic Systems Science; Fukushima University; Fukushima Japan
4. Kansai Research Center; Forestry and Forest Products Research Institute; Kyoto Japan
5. Institute of Boninology; Tokyo Japan