1. Centre d'Ecologie Fonctionnelle et Evolutive CEFE UMR 5175; CNRS-Université de Montpellier-Université Paul-Valéry Montpellier-EPHE; 1919 route de Mende, 34293 Montpellier cedex 5 France
2. Department of Zoology; University of Cambridge; Cambridge CB2 3EJ United Kingdom
3. École Normale Supérieure; 45 rue d'Ulm F-75230 Paris France
4. Integrative Systems Biology Laboratory; King Abdullah University of Science and Technology (KAUST); Thuwal 23955-6900 Kingdom of Saudi Arabia