1. Centre for Integrative Ecology; School of Life and Environmental Sciences; Deakin University; Burwood Victoria 3125 Australia
2. School of Biological Sciences; Monash University; Victoria 3168 Australia
3. Division of Ecology, Evolution and Genetics; The Australian National University; Canberra ACT 0200 Australia
4. School of BioSciences; The University of Melbourne; Victoria 3010 Australia