1. Department of Ecology; Evolution, and Organismal Biology; Iowa State University; Ames IA, 50011 USA
2. Department of Botany; University of Wisconsin Madison; WI, 53706 USA
3. Department of Statistics; University of Wisconsin Madison; WI, 53706 USA
4. Morris Arboretum of the University of Pennsylvania; 100 E. Northwestern Avenue Philadelphia PA, 19118 USA
5. Institute of Fundamental Sciences; Massey University; Palmerston North 4442 New Zealand
6. School of Natural and Environmental Sciences; Newcastle University; NE1 7RU UK
7. Wisconsin Institute for Discovery; 330 N Orchard St Madison WI 53715 USA