1. Department of Biological and Environmental Science; University of Jyväskylä; PO Box 35 FI-40014 Jyväskylä Finland
2. Department of Biological and Environmental Science; Centre of Excellence in Biological Interactions; University of Jyväskylä; PO Box 35 FI-40014 Jyväskylä Finland
3. Finnish Museum of Natural History; Zoology Unit; University of Helsinki; P.O. Box 17 FI-00014 Helsinki Finland