1. Center for International Collaboration and Advanced Studies in Primatology Kyoto University Primate Research Institute Inuyama Aichi 484–8506 Japan
2. Negaunee Integrative Research and Gantz Family Collections Centers Field Museum of Natural History Chicago IL 60605 USA
3. Museums of Paleontology and Vertebrate Zoology University of California Berkeley CA 94720 USA
4. Department of Integrative Biology University of California Berkeley CA 94720 USA