1. Geography Research Unit; University of Oulu; Oulu Finland
2. Finnish Environment Institute; Biodiversity Centre; Oulu Finland
3. Department of Ecology (ICB); Universidade Federal de Goiás; Goiânia GO Brazil
4. Department of Ecosystem Research; Leibniz-Institute of Freshwater Ecology and Inland Fisheries (IGB); Berlin Germany
5. Department of Biological and Environmental Science; University of Jyväskylä; Jyväskylä Finland
6. Department of Geosciences and Geography; University of Helsinki; Helsinki Finland