1. Institut de Recherche pour le Développement; UMR ENTROPIE (IRD-Université de la Réunion-CNRS); BPA5 98848 Nouméa New Caledonia France
2. Department of Biodiversity and Conservation Biology; University of the Western Cape; Bellville 7535 South Africa
3. Institute of Marine Biology; National Taiwan Ocean University; Keelung 20224 Taiwan
4. Dip.to di Bioscienze; Università degli Studi di Milano; via G. Celoria 26 20133 Milan Italy