1. School of Agronomy; Anhui Agricultural University; Hefei 230036 China
2. Key Laboratory of Wheat Biology and Genetic Improvement on South Yellow & Huai River Valley; Ministry of Agriculture; Hefei 230036 China
3. National United Engineering Laboratory for Crop Stress Resistance Breeding; Anhui Agricultural University; Hefei 230036 China
4. Anhui Key Laboratory of Crop Biology; Hefei 230036 China
5. Department of Plant Breeding and Genetics; University of Agriculture Peshawar; Peshawar Pakistan