1. Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany; Chinese Academy of Sciences; Kunming 650201 China
2. University of Chinese Academy of Sciences; Beijing 100049 China
3. Department for Systematic and Evolutionary Botany; University of Vienna; A-1030 Rennweg 14 Austria
4. Key Laboratory of Plant Resources Conservation and Utilization; College of Biology and Environmental Sciences; Jishou University; Jishou Hunan 416000 China