1. Evolution and Ecology Research Centre; School of Biological, Earth and Environmental Sciences; University of New South Wales; Sydney NSW 2052 Australia
2. Sydney Institute of Marine Science; Mosman NSW 2028 Australia
3. Oceans and Atmosphere Flagship; Commonwealth Scientific and Industrial Research Organisation; GPO Box 1538 Hobart TAS 7001 Australia
4. New South Wales Department of Primary Industries; Port Stephens Fisheries Institute; Locked Bag 1 Nelson Bay NSW 2315 Australia
5. WildFish Research; Grays Point NSW 2232 Australia