1. Graduate School of Biosphere Science; Hiroshima University; Kagamiyama 1-7-1, Higashi- Hiroshima 739-8521 Japan
2. Department of Crop Science, Faculty of Agriculture; University of Peradeniya; Peradeniya 20400 Sri Lanka
3. Department of Biology and Department of Biomedical and Molecular Sciences; Queen's University; Kingston Ontario K7L3N6 Canada
4. School of Biological Sciences and Institute of Agriculture; The University of Western Australia; 35 Stirling Highway Crawley (Perth) Western Australia 6009 Australia
5. Max-Planck-Institute for Plant Breeding Research; Carl-von-Linné-Weg 10 50829 Cologne Germany