1. College of Oceanography Institute of Marine Biology Hohai University No.1 Xikang Road Nanjing 210098 China
2. Yantai Institute of Coastal Zone Research Chinese Academy of Sciences 17 Chunhui Road Yantai 264003 China
3. Key Laboratory of Sustainable Utilization of Technology Research for Fishery Resource of Zhejiang Province Marine Fisheries Research Institute of Zhejiang Province Zhoushan 316021 China