1. Department of Agricultural Biotechnology Seoul National University Seoul 08826 Korea
2. Department of Biology Ajou University Suwon 16499 Korea
3. Plant Genomics and Breeding Institute Seoul National University Seoul 08826 Korea
4. Graduate School of Science and Technology, Nara Institute of Science and Technology, Ikoma Nara 630‐0192 Japan
5. Department of Biology University of Washington Seattle WA 98195 USA
6. Institute of Agricultural Life Sciences Seoul National University Seoul 08826 Korea