1. Research Institute for Humanity and Nature 457‐4 Motoyama, Kamigamo, Kita‐ku Kyoto 603‐8047 Japan
2. Japan Agency for Marine‐Earth Science and Technology 2‐15 Natsushima‐Cho Yokosuka Kanagawa 237‐0061 Japan
3. Graduate School of Life Sciences Tohoku University 6‐3 Aramaki‐Aoba, Aoba‐ku Sendai Miyagi 980‐8578 Japan
4. Tohoku National Fisheries Research Institute Japan Fisheries Research and Education Agency 3‐27‐5, Shinhama‐cho Shiogama Miyagi 985‐0001 Japan
5. Graduate School of Fisheries Sciences Hokkaido University 3‐1‐1 Minatomachi Hakodate Hokkaido 041‐8611 Japan
6. Atmosphere and Ocean Research Institute The University of Tokyo 5‐1‐5, Kashiwanoha, Kashiwa‐shi Chiba 277‐8564 Japan
7. Hokkaido National Fisheries Research Institute Japan Fisheries Research and Education Agency 2‐2 Nakanoshima, Toyohira‐ku Sapporo Hokkaido 062‐0922 Japan