1. Pacific Salmon Ecology and Conservation Laboratory, Department of Forest and Conservation Sciences, The University of British Columbia, Vancouver, BC V6T 1Z4, Canada.
2. Pacific Salmon Foundation, Vancouver, BC V6J 4S6, Canada.
3. Kintama Research Services, Ltd., Nanaimo, BC V9V 1N8, Canada.
4. Department of Biological Sciences, University of New Hampshire, Durham, NH 03824, USA.
5. Fisheries and Oceans Canada, West Vancouver Laboratory, West Vancouver, BC V7V 1H2, Canada.