1. Centre for Fisheries Ecosystems Research, Fisheries & Marine Institute, Memorial University of Newfoundland, St John's NL A1C 5R3, Canada
2. Fisheries and Oceans Canada, 100 Annacis Pkwy, Delta, BC V3M 6A2, Canada
3. College of Marine Science, Shanghai Ocean University, 999 Huchenghuan Road, Shanghai 201306, China