1. Oregon Cooperative Fish and Wildlife Research Unit, Department of Fisheries, Wildlife and Conservation Sciences, Oregon State University, Corvallis, OR 97330, USA
2. U.S.D.A. Forest Service, Pacific Northwest Research Station, Olympia, WA 98512, USA
3. U.S. Geological Survey, Oregon Cooperative Fish and Wildlife Research Unit, Department of Fisheries, Wildlife and Conservation Sciences, Oregon State University, Corvallis, OR 97330, USA