1. Department of Watershed Science, Utah State University, 5200 Old Main Hill, Logan, UT 84322, USA; The Ecology Center, Utah State University, 5205 Old Main Hill, Logan, UT 84322, USA.
2. Department of Wildland Science, Utah State University, 5200 Old Main Hill, Logan, UT 84322, USA; The Ecology Center, Utah State University, 5205 Old Main Hill, Logan, UT 84322, USA.
3. US Geological Survey, Utah Cooperative Fish and Wildlife Research Unit, Department of Watershed Sciences, Utah State University, 5210 Old Main Hill, Logan, UT 84322, USA; The Ecology Center, Utah State University, 5205 Old Main Hill, Logan, UT 84322, USA.
4. Department of Watershed Science, Utah State University, 5200 Old Main Hill, Logan, UT 84322, USA.
5. US Bureau of Reclamation, 125 State Street #6103, Salt Lake City, UT 84138, USA.