1. B.C. Ministry of Environment, Conservation Science Section, 2202 Main Mall, The University of British Columbia, Vancouver, BC V6T 1Z4, Canada; Institute for the Oceans and Fisheries, The University of British Columbia, 2202 Main Mall, Vancouver, BC V6T 1Z4, Canada.
2. The University of British Columbia, Department of Zoology, 6270 University Boulevard, Vancouver, BC V6T 1Z4, Canada.