1. University of Toronto Mississauga, Department of Biology, Mississauga, Ontario, Canada;
2. University of Toronto, Department of Ecology and Evolutionary Biology, Toronto, Ontario, Canada
3. Ontario Ministry of Natural Resources, Aquatic Ecosystem Science Section, Peterborough, Ontario, Canada;
4. Ontario Ministry of Natural Resources, Maple, Ontario, Canada, ;
5. University of Toronto - Mississauga, 71637, Biology, Mississauga, Ontario, Canada;