1. Department of Biological and Environmental Sciences, University of Gothenburg, Box 463, SE-405 30, Gothenburg, Sweden.
2. The Key Laboratory of Mariculture, Ministry of Education, Ocean University of China, Qingdao 266003, China.
3. Laboratoire Evolution et Diversité Biologique (EDB UMR 5174), Université de Toulouse, CNRS, ENFA, UPS, 118 route de Narbonne, F-31062 Toulouse, France.