1. Department of Biology, Norwegian University of Science and Technology (NTNU), Trondheim, Norway.
2. Norwegian Institute for Nature Research (NINA), Trondheim, Norway.
3. Department of Natural History, Norwegian University of Science and Technology (NTNU), Trondheim, Norway.
4. Centre for Biodiversity Dynamics, Department of Biology, Norwegian University of Science and Technology (NTNU), Trondheim, Norway.