1. School of Aquatic and Fishery Sciences, University of Washington, Seattle, WA, USA
2. Washington Department of Fish & Wildlife, Olympia, WA, USA
3. Auke Bay Laboratories, Alaska Fisheries Science Center, NMFS, NOAA, Seattle, WA, USA
4. Resource Ecology and Fisheries Management Division, Alaska Fisheries Science Center, NMFS, NOAA, Seattle, WA, USA
5. Habitat and Ecological Processes Research Program, Alaska Fisheries Science Center, NMFS, NOAA, Seattle, WA, USA