1. UMR ESE Ecology and Ecosystem Health, Institut Agro, INRAE, 35042 Rennes, France.
2. MIAME, Management of Diadromous Fishes in their Environment, OFB, INRAE, Institut Agro, UPPA-E2S.
3. INRAE, pôle OFB–INRAE–Institut Agro–UPPA MIAME – Management of Diadromous Fish in their Environment, U3E, F-35042, Rennes, France.
4. Salmon and Trout Research Centre, Game and Wildlife Conservation Trust, FBA River Laboratory, Wareham, Dorset, BH20 6BB, UK.