1. Faculty of Biosciences, Fisheries, and Economics, University of Tromsø, N-9037 Tromsø, Norway.
2. Museum of Natural History and Archaeology, Norwegian University of Science and Technology (NTNU), N-7491 Trondheim, Norway.
3. Faculty of Biosciences, Fisheries, and Economics, University of Tromsø, N-9037 Tromsø, Norway; Norwegian Institute for Nature Research, N-7485 Trondheim, Norway.
4. Graduate School of Informatics, Kyoto University, Kyoto 606-8501, Japan.
5. Institute of Marine Research, N-9294 Tromsø, Norway.
6. Norwegian Institute for Nature Research, N-7485 Trondheim, Norway.