1. Department of Ecology and Evolutionary Biology, University of Toronto, 25 Willcocks Street, Toronto, ON M5S 3B2, Canada; Department of Biology, University of Toronto at Mississauga, William G. Davis Building, 1867 Inner Circle Road, Mississauga, ON L5L 1C6, Canada.
2. Department of Biology, University of Toronto, William G. Davis Building, 1867 Inner Circle Road, Mississauga, ON L5L 1C6, Canada.
3. Aquatic Research and Monitoring Section, Ontario Ministry of Natural Resources and Forestry, 2140 East Bank Drive, Peterborough, ON K9J 7B8, Canada.