1. US Fish and Wildlife Service, Columbia River Fish and Wildlife Conservation Office, 1211 SE Cardinal Court, Suite 100, Vancouver, WA 98683, USA.
2. Oregon Cooperative Fish and Wildlife Research Unit, US Geological Survey, Department of Fisheries and Wildlife, Oregon State University, 104 Nash Hall, Corvallis, OR 97331-3803, USA.
3. US Geological Survey, Forest and Rangeland Ecosystem Science Center, 3200 Southwest Jefferson Way, Corvallis, OR 97331, USA.
4. US Fish and Wildlife Service, Fish and Aquatic Conservation Program, Pacific Regional Office, 911 NE 1st Avenue, Portland 97232, USA.