1. Institute of Biology, University of Neuchâtel, 11, rue Emile-Argand, 2009 Neuchâtel, Switzerland.
2. Department of Biological Sciences, University of Calgary, 2500 University Drive Northwest, Calgary, AB T2N 1N4, Canada.
3. Department of Zoology, University of Cambridge, Downing Street, CB2 E3J, Cambridge, UK.
4. Department of Ecology and Evolutionary Biology, University of Arizona, P.O. Box 210088, Tucson, AZ 85721, USA.