1. Faculty of Land and Food Systems, University of British Columbia, Vancouver, BC V6T 1Z4, Canada.
2. Faculty of Forestry, University of British Columbia, Vancouver, BC V6T 1Z4, Canada.
3. BC Ministry of Forests, Lands and Natural Resource Operations and Rural Development, Kamloops, BC V2C 2T3, Canada.
4. Department of Botany, University of British Columbia, Vancouver, BC V6T 1Z4, Canada.