1. Centre for Aquatic Environments, School of Environment and Technology, University of Brighton, Brighton BN2 4GJ, UK.
2. School of Geography, Archaeology and Environmental Studies, University of the Witwatersrand, Private Bag 3, Wits 2050, South Africa.
3. National Oceanography Centre (Southampton), School of Ocean and Earth Science, University of Southampton, Southampton SO14 3ZH, UK.