1. Department of Wildlife Management, Sokoine University of Agriculture (SUA), P.O. Box 3073, Morogoro, Tanzania
2. African Centre of Excellence for Innovative Rodent Pest Management and Biosensor Technology Development, SUA, Morogoro, Tanzania
3. Department of Veterinary Anatomy and Pathology, Sokoine University of Agriculture, P.O. Box 3000, Morogoro, Tanzania
4. Mammal Research Institute, Department of Zoology & Entomology, University of Pretoria, Pretoria 0002, South Africa