1. Department of Integrative Biology, University of Guelph, ON, Canada
2. Office of the Deputy Vice-Chancellor: Research, Innovation and Engagement, Tshwane University of Technology, Pretoria, South Africa
3. National Museum of Namibia, P.O. Box 147, Windhoek, Namibia
4. Unit for Environmental Sciences and Management, North-West University, Potchefstroom, South Africa